After facing off with the Mogadorian ruler and almost being annihilated, the Garde go into hiding. They must learn to master their Legacies and learn to work together as a team. When they receive a sign from Number Five -- a crop circle in the shape of a Loric symbol -- they know they are close to being reunited. But could it be a trap? Time is running out, and the only thing they know for certain is that they have to get to Five before it's too late....

In a Spanish convent, seventeen-year-old Marina longs to join forces with her fellow Loriens to prepare for battle with the Mogadorians who destroyed their home planet, while in the United States, John, Six, and Sam elude authorities who think John is a terrorist.

In order to thwart the Mogadorian invasion, a group of the Garde head to a pyramid in Mexico in order to awaken a power that can save the world.